Ephemeroptera: Heptageniidae of Gunnison County, ColoradoIntroduction to Heptagenia
Pale Evening Dun, Clinger MayflyWalsh, 1863
Updated 28 September 2025
TSN 100602
Species List
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References
Argyle,DW; Edmunds,GF 1962 Mayflies (Ephemeroptera) of the Curecanti Reservoir Basins Gunnison River, Colorado. University of Utah Anthropological Papers 59 (8) 178-189.
Bednarik,AF; Edmunds,GF 1980 Descriptions of larval Heptagenia from the Rocky Mountain region (Ephemeroptera: Heptageniidae). Pan-Pacific Entomologist 56, 51-62. PDF
Edmunds Jr,GF and McCafferty,WP 1988 The mayfly subimago. Annual review of entomology, 33(1)509-527. PDF
Quote from page 522: "The speed with which subimagos were able to right themselves after being
placed on their side with wings contacting water varied immensely among subimagos of different genera observed (G. F. Edmunds,Jr. and T. J. Fink,
unpublished). The results could be based on differences in hydrofuge capacity, differences in behavioral response among mayflies, or both. Siphlonurids (Siphlonurus, Ameletus, Parameletus) were slow to respond, and most of the individuals, especially of Siphlonurus, were entrapped on the surface. Heptageniids (Stenacron, Nixe, Heptagenia) righted themselves faster than the siphlonurids and almost always escaped successfully. Leptophlebiids (Paraleptophlebia) and ephemerellids (Ephemerella, Drunella) responded rapidly and escaped so fast that sometimes it was impossible to see if they righted themselves before taking flight"
Flowers,RW 1980 A review of the Nearctic Heptagenia (Heptageniidae, Ephemeroptera). Pages 93-102 in Flannagan JF; Marshall KE (eds.), 1980, Advances in Ephemeroptera Biology. Plenum Press, New York. PDF
Flowers,RW 1980 Two new genera of Nearctic Heptageniidae (Ephemeroptera). Florida Entomologist 63:296-307. PDF
Splits the new genera Leucrocuta and Nixe from Heptagenia.
Walsh,BD 1863 Observations on certain N. A. Neuroptera, by H. Hagen, M.D., of Koenigsberg, Prussia; translated from the original French ms., and published by permission of the author, with notes and descriptions of about twenty new N.A. species of Pseudoneuroptera. Proceedings of the Entomological Society of Philadelphia 2:167–272.
Webb,JM and McCafferty,WP 2008 Heptageniidae of the world. Part II. Key to the genera. Canadian Journal of Arthropod Identification, 7(10.37551). PDF
Webb,J; McCafferty,W. and Lehmkuhl,D 2002 The larval stage of Heptagenia adaequata. Entomological News, 113(5), pp.289-293. PDF
Abstract: "Larvae of the poorly known western North American species Heptagenia adaequata are described for the first time based on material reared from the Saskatchewan River system. Diagnoses are provided for differentiating larvae of H. adaequata from other species of Heptagenia within its range, especially the common H. solitaria and H. elegantula, with which it might be confused."
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